Showing posts with label Gokturks. Show all posts
Showing posts with label Gokturks. Show all posts

Friday, November 15, 2024

Another step forward in solving the puzzle of the Pannonian Avars

Over the last two years the Avar period has probably become the most extensively covered historical period when it comes to the field of ancient DNA. While I welcome all forms of information regarding steppe nomads, it does make me wonder why precisely this period out of all phases of steppe nomad history or European history gets to enjoy such a degree of dense sampling. Granted, the huge number of samples we have come from Network of large pedigrees reveals social practices of Avar communities where multiple generations of several Avar period family cemeteries were sequenced. 

I guess the team at HistoGenes felt like adding fuel to the flame, throwing 722 more individuals onto the pile for an upcoming DNA article! The article has not been released yet but the project data has been uploaded:

Project: PRJEB76548

After a long-distance migration from the East, Avar people of Eastern Asian ancestry arrived in Eastern Central Europe in 567/568 CE and encountered groups with very different, European ancestry1,2. We used ancient genome-wide data of 722 individuals and fine-grained interdisciplinary analysis of large 7th-8th c. CE neighboring cemeteries south of Vienna (Austria) to address the centuries-long impact of this encounter1,2. We found that the ancestry at one site (Leobersdorf) was and remained dominantly East-Asian-like, even 200 years after the Avar immigration, while the other site (Mödling) displays local, European-like ancestry. These two nearby sites exhibit very few links of direct biological relatedness, despite sharing a distinctive late-Avar culture3,4. We reconstructed 6-generation pedigrees at both sites linking together up to 450 closely-related individuals, allowing per-generation demographic profiling of the communities. Despite different ancestry, these pedigrees together with large networks of distant relatedness display an absence of consanguinity, a strong patrilineal pattern with female exogamy, multiple reproductive partnerships (e.g. levirate) and direct correlation of social status to biological connectivity through markers of high social status in the archaeological material. The generation-long genetic barrier was maintained by systematically choosing partners with similar ancestry from other sites in the Avar realm. Leobersdorf had more biological connections with the Avar heartlands than Mödling, which is instead linked to another site from the Vienna Basin with European-like ancestry. Mobility between sites was mostly due to female exogamy pointing to different marriage networks as the main driver of the maintenance of the genetic barrier.

So yeah another upcoming article with hundreds of Avar period samples, most from family graves. What’s the big deal then? We’ve seen it already. Avars were of East Asian origin, and are genetically similar to Xianbei and Rouran period individuals from Mongolia and China. These Avars seem predominantly to be carriers of haplogroup N of East Asian origins. Amongst these Avars there also is a presence of Oghuric Turkic nomads, and we see various European populations living within the Avar realm. What more interesting can be found from studying these two cemeteries?


A new paternal lineage

About two years ago I wrote a post about the Pannonian Avars when our first DNA samples of the population were published. It ended with the conclusion that I thought these genomic profiles were indicative of the Avars likely being of (Para-)Mongolic origin given their strong  (Para-)Mongolic signatures.


But there always was a doubt regarding that attribution since we only had one Y-chromosome lineage we could connect to the core Avar population, N-F4205, downstream of haplogroup N-L1026.  Aside from the litany of N-L1026 in Uralic populations, the specific N-F4205 lineage shows up as early as the Xiongnu period withsample IMA005 from Buryatia.


This time though? The situation seems a bit different. The Avar period samples at Leobersdorf seem to primarily carry C2-M217 rather than the common N-F2405 lineage of the Avars. The more detailed subclade seems to be C-F3830, although it is not certain if all samples had this subclade or perhaps other forms of C-M217.


You can find a whole lot of haplogroup calls for the new Avar samples in this google sheet file: 2024_Wang_ViennaBasinAvarPeriod. I am not the author by the way, and I currently do not know who actually made this file. Will the Real Slim Shady please stand up?


The Leobersdorf cemetery has revealed something that all the previous samples of the Avars failed to show, a genetic connection that unquestionably ties the pannonian Avars to their contemporary Mongolic populations. C-F3830 is a Y-chromosome haplogroup which consistently shows up in ancient populations connected to the Donghu, and it is undoubtedly one of the main haplogroups of the Mongolic peoples. This is interesting because C-F3830 is a subclade of C-F1756, and other C-F1756 clades such as C-Y10420 seem to be very significant for Proto-Turkic people, showcasing a paternal relation between their respective founding populations.

The Donghu (meaning Eastern Barbarians) were a historical nomadic confederation situated in modern day Inner Mongolia and Liaoning, China. After their defeat by the Xiongnu, the Donghu split into two factions; The Xianbei and the Wuhuan. From these two factions (primarily the Xianbei though) we get the Tuoba, Rouran, Khitans and the Mongols. The Pannonian Avars can now be added to the list as well thanks to the samples from the Leobersdorf cemetery. But who were the people at the Leobersdorf cemetery?


Leobersdorf Avar cemetery

The Leobersdorf cemetery offers a fascinating look at Avar society just after their arrival in the Vienna Basin in the late 6th century. On the Thanados website you can actually find quite a bit of information from the burials at the Leobersdorf site. I’m not sure if every single burial has been featured but for most of these you can actually find quite a bit of information about the grave sizes and contents. For example, from grave 071 (which I picked randomly) I was able to find an image of the skeleton layout and some of the burial goods such as belt fittings and a sabre.


The graves themselves are spaced differently depending on their period and style. The early “scattered” graves  sit a bit apart from the later rows, which suggests a shift from individual or small-group burials to a more standardised cemetery layout over time. As more Avars settled in the area, the cemetery expanded to include around 153 graves in organised rows, with burials spanning from the 7th to 8th centuries [1]. 


Leobersdorf’s burials also offer clues about daily life and social roles within this Avar community. Unlike some Avar cemeteries where warrior items dominate, Leobersdorf contains relatively few overtly martial artefacts, with most grave goods leaning toward personal ornamentation like earrings, belt fittings, and the occasional gilded buckle. This might suggest that while status was marked through adornment, the site wasn't a frontline warrior society. There are few indicators of violence in the skeletal remains, and injuries are largely limited to healed fractures, hinting at a community life that wasn’t defined by constant conflict.

Interestingly, there also is a clear pattern in the burial treatment of males and females: men often have more elaborate belt fittings, perhaps reflecting roles tied to their status, while women’s graves tend to include decorative jewellery. The Leobersdorf graves do not show the splendor you would find in some of the elite Avar burials of the Carpathian Basin, no piles of gold or elaborate weaponry. But that does not mean the burials were modest either. The cemetery includes belt fittings, buckles, earrings and gilded bronze items, indicating that social rank mattered here too. Men’s graves often have complex belt sets with animal motifs, possibly marking positions within the community.

One of the earliest burials at Leobersdorf, dating to the 6th century AD, is quite intriguing as the burial goods contained a silver burial mask and rosette fittings from a bow quiver. That sounds like a pretty unusual set of items, especially for the time, hinting that whoever was buried there might have had status or connections. Similar burial customs with masks would later appear in Khitan burials, though they come centuries after this one and we should not assume a direct link.

Two burial masks of the Liao dynasty period
Rouran connection

The presence of C-F1380 in Avar populations in Europe is highly interesting, but this does not change the stance I had regarding the Avar khaganate and the Rouran Khaganate connection in my previous blog post.


It is important to recognize that the N-F4205 lineage was found in the most elite of Avar burials, but also the most simple burials belonging to commoners. Although we do not have burials which can be connected to the Avar Khagans with certainty, the richest burial of the early Avar period belonged to sample A1802 which had N-F4205 [2]. In order to prove the Rouran-Avar connection you’d have to show strong evidence that the rulers of the Rouran Khaganate also had this clade.


It is interesting to note that two years after this post we received confirmation that Emperor Wu of the northern Zhou dynasty, of Xianbei origin, carried C-F380 > C-Z603 in the article Ancient genome of the Chinese Emperor Wu of Northern Zhou by Du et al. If you want to play around with the genome of Emperor Wu on Vahaduo, you can access the G25 coordinates here.

Funnily enough according to the Book of Wei this tribe had an origin in the Xiongnu and later joined the Xianbei. Either this record is not correct (the book of Wei is known to be less accurate than other books) or that southern Xiongnu clan was of Donghu origin - which is possible but would be one hell of a coincidence.


Although the lineage of the Northern Zhou dynasty in no way guarantees that the rulers of the Rouran Khaganate also carried C-F380, it does underline the importance of C-F380 in early Mongolic populations. Considering the origin of the Mugulü clan, being intrinsically tied to the steppe regions adjacent to the Gobi Desert, I think there is a really high chance C-F380 would be their patrilineal lineage.


Varchonites and Pseudo-Avars

Back to the Avars. It would be interesting to figure out if the C-F380 Avars were of the same population/ethnic group as the N-F4205 Avars, or if they represent two different communities within the Avars. Perhaps it is a reflection of the ethnic situation as described by Theophylact Simocatta? It was said that the Avars were composed of two groups, the Var and Chunni (from which we get Varchonites). 


Here is a section from Theophylact’s writings (emphasis is mine):

But since we have made reference to the Scythians, both those in the Caucasus and those who face northwards, come then, let us interrupt our history and present, like an intercalated narrative, the events which attended these very great nations during these times. When summer had arrived in this particular year, he who is celebrated by the Turks as Chagan in the east dispatched ambassadors to the emperor Maurice; he composed a letter and inscribed in it victory-praises.


The letter's salutation was as follows, word for word: ‘To the king of the Romans, the Chagan, the great lord of seven races and master of seven zones of the world.’ For this very Chagan had in fact outfought the leader of the nation of the Abdeli (I mean indeed, of the Hephthalites, as they are called), conquered him, and assumed the rule of the nation.


Then he was greatly elated at the victory and, making an alliance with Stembischagan, he enslaved the Avar nation. But let no one think that we are distorting the history of these times because he supposes that the Avars are those barbarians neighbouring on Europe and Pannonia, and that their arrival was prior to the times of the emperor Maurice. For it is by a misnomer that the barbarians on the Ister have assumed the appellation of Avars; the origin of their race will shortly be revealed. 


So, when the Avars had been defeated (for we are returning to the account), some of them made their escape to those who inhabit Taugast. Taugast is a famous city, which is a total of one thousand five hundred miles distant from those who are called Turks, and which borders on the Indians. The barbarians whose abode is near Taugast are a very brave and numerous nation, and without rival in size among the nations of the world. Others of the Avars, who declined to humbler fortune because of their defeat, came to those who are called Mucri; this nation is the closest neighbour to the men of Taugast; it has great might in battle both because of its daily practice of drill and because of endurance of spirit in danger. 


Then the Chagan embarked on yet another enterprise, and subdued all the Ogur, which is one of the strongest tribes on account of its large population and its armed training for war. These make their habitations in the east, by the course of the river Til, which Turks are accustomed to call Melas. The earliest leaders of this nation were named Var and Chunni; from them, some parts of those nations were also accorded their nomenclature, being called Var and Chunni. 


Then, while the emperor Justinian was in possession of the royal power, a small section of these Var and Chunni fled from that ancestral tribe and settled in Europe. These named themselves Avars and glorified their leader with the appellation of Chagan. Let us declare, without departing in the least from the truth, how the means of changing their name came to them. When the Barselt, Onogurs, Sabir, and other Hun nations in addition to these saw that a section of those who were still Var and Chunni had fled to their regions, they plunged into extreme panic, since they suspected that the settlers were Avars. For this reason, they honoured the fugitives with splendid gifts and supposed that they received from them security in exchange. 


Then, after the Var and Chunni saw the well-omened beginning to their flight, they appropriated the ambassadors' error and named themselves Avars: for among the Scythian nations that of the Avars is said to be the most adept tribe. In point of fact, even up to our present times, the Pseudo-Avars (for it is more correct to refer to them thus) are divided in their ancestry, some bearing the time-honoured name of Var while others are called Chunni.”

This text opens a whole can of worms, as it is this text that more or less spawned all the debate about the Avar origin and migration. After all, it is the mention of the defeat of the Avars and them fleeing to Taugast (China) that echoes the defeat of the Rouran by the Göktürks.


According to Theophylact (relying on Menander as well as some unknown documents) Türk envoys had conveyed the message that the Pannonian Avars were not the real Avars but Varchonites, mistaken to be Avars by the Onogurs and Sabirs who were attacked by them.


The Göktürks heavily disliked the idea that the Eastern Roman empire had befriended the Avars. Ultimately the problem lies in that while you can have many Khans, there can be only one Khagan according to the Celestial Turks. With the Göktürks and the Avars you have two entities claiming this title at the same time, which was a problem of course. This ultimately is the motivation for calling the Pannonian Avars Pseudo-Avars and Varchonites. Delegitimizing their existence as Avars, and delegitimizing their claim to a title such as Khagan [3]. It is important to point out that the Pannonian Avars have only referred to themselves as Avar, never as Varchonite.


Several authors seem to suggest that Theoplylact had probably combined messages from different periods. Harmatta laid out a convincing argument that the discussion of the subjugation of the Avars does not refer to the defeat of the Rouran at all, but rather refers to military conflicts in the final part of the 6th century AD. The “Avars”  discussed could actually be a Tiele tribe called Apar or Abar [4].


I do wonder though, if the Rouran and "Asiatic Avars" were distinct, why would there be an effort to declare the Pannonian Avars as fraudulent Varchonites who falsely took on the name Avar? The motivation ultimately seems to be that they called themselves Khagan, a title once held by the Rouran. Perhaps this had nothing to do with the Rouran per se, but was simply just an attempt to delegitimize the Pannonian Avars and their claim to the Khagan title.


I also  wonder what to make of the Avars (Ἄβαροι) mentioned in the fragments of Priscus:

At this time the Saraguri, Urogi and the Onoguri sent envoys to the eastern Romans. These tribes had left their native lands when the Sabiri attacked them. The latter had been driven out by the Avars who had in turn been displaced by the tribes who lived by the shore of the Ocean. In the same way, the Saraguri, driven to search for land, came into contact with the Akatirian Huns and, after engaging them in many battles, defeated that tribe. The Saraguri then approached the Romans, wishing to win their friendship, and the Emperor and his courtiers received them in a kindly manner, gave them gifts and sent them away.

These Avars drove out the Sabinores having themselves been displaced by the tribes who lived by the shore of the Ocean. The latter had left their land on account of the mist which came from an inundation of the Ocean and because a flock of gryphons had appeared. It is said that they would not leave until they had eaten the whole race of men. Therefore, driven out by these evils, the ocean dwellers fell upon their neighbours, and since the attackers were more powerful, the Avars, who could not resist their onset, were displaced. In the same way the Saraguri were driven out and came into contact with the Akatirian Huns.

Were these Avars mentioned by Priscus the Rouran, an early attestation of the Pannonian Avars, or a completely different tribe which just happened to have a similar name?


The origins of the Avars from a historical perspective is a highly complicated puzzle, and attempting to solve it through historical sources is a new notches above my pay grade. I mean, for over 2 centuries historians have attempted this without any consensus forming to this day, so it would be unlikely that Musaeum Scythia is going to have the answers.


How can DNA help solve this historical puzzle?

However, we can use DNA of course. The DNA of the Pannonian Avars really points to a population migrating from the Mongolian plateau to Eastern Europe directly, without having a presence of several centuries in Central Asia. 


By default this would make it hard to connect the Pannonian Avars to a migration out of the Hephthalite realm. The likelihood that this population was of Mongolic origin (although we cannot know if they still spoke Mongolic) is also high, and Varchonites would more likely refer to Oghuric tribes in Central Asia and Eastern Europe.


Although related to this it is perhaps interesting to bring up DA95 again. This sample had the Avar N-F4205 lineage and a similar autosomal profile, in my mind there undoubtedly was a connection. But this sample was buried in Krasnodar, Kazakhstan while dating to 331- 354 AD [5]. Could this individual be one of the Avars mentioned by Priscus? Given how it precedes the source by a century I am not too sure, unless the events recorded by Priscus preceded him by a century?


The C-F380 lineage in this population provides a closer tie to the Xianbei-Wuhuan core population, including the Rouran. However the vast majority of Avars, including elites had a different lineage. I still doubt that the profile and paternal lineages of the early Pannonian Avars can be connected to the rulers of the Rouran khaganate for the same reason as I outlined some years ago.


The Heqin policy should have had an effect on their genomes and we see none of that in both the elites and commoners of the Pannonian Avars. If we take Wu of Northern Zhou, whose family were not derived from the rulers of the Tuoba Wei, he had at least one Han grandmother and about 30% Yellow River related ancestry, suggesting additional admixture events with people from the Central Plains. 


I still think this population could have represented a powerful clan from the northern part of the Rouran khaganate. The Baikal region perhaps? It is pretty close to the location of Xiongnu sample IMA005 with the N-F4205 lineage. The N-F4205 lineage also exists in modern Buryats which could point to this direction.


This could explain the wealth and power this population had, while also explaining why their genetic profile plots so northerly and did not partake in Heqin policies. Maybe their ancestors were part of the Xiongnu empire a long time ago but were assimilated into Mongolic communities during the Xianbei or early Rouran periods. 


It would be very interesting if it turns out that the C-F380 lineage could be connected to the rulers of the Rouran however. Perhaps it could be that the C-F380 lineage with the collapse of the Rouran Khaganate joined up with the N-L1026 clan carrying the prestige and Khagan title with them. Sometime in that process, The N-L1026 clan overtakes the C-F380 clan in terms of political power and takes the Khagan title. 


Imagine how much of a twist that would be? From the Avar perspective their usage of Khagan would be perfectly legitimate, whereas the viewpoint of the Göktürks that they were not the real Khagans was also justified.


References:

  1. Daim, F. (2020), Awarische Gräberfelder zwischen Petronell und Leobersdorf – Neue archäologische und anthropologische Untersuchungen, in Awarische Gräberfelder zwischen Petronell und Leobersdorf: Neue Forschungen zur Archäologie der Awarenzeit in Österreich, Propylaeum.
  2. Gnecchi-Ruscone, G. A., Szécsényi-Nagy, A., Koncz, I., Csiky, G., Rácz, Z., Rohrlach, A. B., Brandt, G., Rohland, N., Csáky, V., Cheronet, O., Szeifert, B., Rácz, T. Á., Benedek, A., Bernert, Z., Berta, N., Czifra, S., Dani, J., Farkas, Z., Hága, T., Hajdu, T., … Krause, J. (2022). Ancient genomes reveal origin and rapid trans-Eurasian migration of 7th century Avar elites. Cell, 185(8), 1402–1413.e21. https://doi.org/10.1016/j.cell.2022.03.007
  3. Mihály Dobrovits (2003), "They Called Themselves Avar" (Webfestschrift Marshak 2003), Transoxiana
  4. János Harmatta (2001), Avar History and Diplomacy in Late Antiquity, Hungarian Academy of Sciences.
  5. Damgaard, P. B., Marchi, N., Rasmussen, S., Peyrot, M., Renaud, G., Korneliussen, T., Moreno-Mayar, J. V., Pedersen, M. W., Goldberg, A., Usmanova, E., Baimukhanov, N., Loman, V., Hedeager, L., Pedersen, A. G., Nielsen, K., Afanasiev, G., Akmatov, K., Aldashev, A., Alpaslan, A., Baimbetov, G., … Willerslev, E. (2018). 137 ancient human genomes from across the Eurasian steppes. Nature, 557(7705), 369–374. https://doi.org/10.1038/s41586-018-0094-2

Tuesday, March 21, 2023

Ancient DNA from a Xiongnu period elite cemetery in western Mongolia and Empress Ashina

 This week is a very interesting one when it comes to access of ancient DNA coming from the eastern steppes. We got a 2-for-1 combo deal with ancient DNA from the Xiongnu period and the Goktürk period.

Ancient DNA from Xiongnu cemeteries in Western Mongolia

The other day raw data for an upcoming article headed by Choongwon Jeong, who also was the lead author of the 2020 article “A Dynamic 6,000-Year Genetic History of Eurasia’s Eastern Steppe” which currently is one of the most significant articles to have come out in relation to the topic of the eastern steppes during the metal ages was uploaded to Edmond (the Open Research Data Repository of the Max Planck Society). 


Description:

This collection includes the EIGENSTRAT format genotype data of 17 ancient individuals for the 1240K SNP panel. The individuals are newly analyzed in the study "Genetic population structure of the Xiongnu Empire at imperial and local scales" and are from two Xiongnu-period archaeological sites in Mongolia: TAK(Takhiltyn Khotgor) and SBB(Shombuuzyn Belchir).

The raw data is accessible here:

Data for "Genetic population structure of the Xiongnu Empire at imperial and local scales" - Edmond


Davidski from the Eurogenes blog has converted these samples to Global25 coordinates, which you can access here.

https://drive.google.com/file/d/1PC-N4dJr3pWynJIYKdAYzZWwXhJug5e-/view?usp=share_link


This means that you can now utilize these samples with the various Vahaduo tools, which you can do here:

https://vahaduo.github.io/


Visit the Eurogenes blog for the Global25 coordinates:

Eurogenes Blog: Getting the most out of the Global25


I will discuss the genetic aspect in a bit, but first I will go over the two sites these samples come from. Below is a map which showcases the two sites, Shombuuzyn Belchir and Takhiltyn Khotgor being site 1 and 2 on the map respectively [1]. 


Takhiltyn Khotgor is an interesting site because it is one of the few Xiongnu elite cemeteries which are present in western Mongolia, as most of the large cemeteries are in Central Mongolia and the Transbaikal region [2].


Now I think that due to the limited information we have about the Xiongnu empire a lot of opinions about “Xiongnu elites” have been pulled right out of thin air. Reading some historian’s works you’d get the impression that it is certain that the Xiongnu elites belonged to a singular ethnic group, whether it’s Yeniseian speaking, Turkic speaking, Iranian speaking. Personally I see no real evidence for this in primary sources, in fact I see no evidence that the founding tribes of the Xiongnu confederation themselves belonged to a single ethnolinguistic group, let alone the elites at an imperial or local level of the imperial period.


Nevertheless getting genomic data from such elite burials will be very informative and could help to confirm or debunk the many many claims that float around about the Xiongnu and the elite strata of their society.


The arrangement and components of tomb complexes at Takhiltyn Khotgor, as well as the

contents of the main tombs, closely resemble elite Xiongnu cemeteries found elsewhere. The

elite graves here are undoubtedly associated with the elite imperial culture of the Xiongnu

polity and thus may be considered within the limits of the empire. Despite the consistencies

in mortuary remains for the elite, evidence arises in the satellite burials of differentiation from

standard Xiongnu customs. Interments in stone cists such as THL-64-1 and THL-64-2 are

seldom found in other Xiongnu cemeteries, and the flexed legs of these two graves and THL25-1 deviate from the prevailing custom of fully stretched supine burials in Xiongnu graves

(Törbat 2004). The most significant difference is the surface demarcations. The small tight

cluster of stones over the grave pit differs greatly from the tradition of stone rings. It is not

clear what these variations indicate, but it may relate to either local variations of the Altai or

perhaps a social group not yet delineated within the greater corpus of Xiongnu remains. Further excavations of Xiongnu graves, especially outside the large elite cemeteries, will need to be

conducted in the area of Khovd aimag in order to further clarify this issue.


Crescent sun disc from grave 64 at Takhiltyn Khotgor


Shombuuzyn Belchir is a bit less extravagant than Takhyltin Khotgor. The burials here generally fall in line with the typical Xiongnu burial traditions of Central and eastern Mongolia.

The radiocarbon dates for these sites range in between 50 BC and 100 AD for Takhyltin Khotgor and Shumbuulzyn Belchir [3].


This is an interesting period during the Xiongnu empire as it falls in the range of the first Xiongnu civil war, where we see Zhizhi Chanyu making his base around the Altai region. Presumably, this was after the defeat of the rebelling Hujie Chanyu, the Hujie being one of the tribes of the Xiongnu most likely located in the southern Altai region. 


Bernshtam interpreted the historical records as indicating that Zhizhi Chanyu made his base around the  “celestial mountain” as Zhizhi making his base around the Tian Shan mountains, and that this migration was the movement of Huns into Central Asia. However, based on the description it seems far more likely that the “celestial mountain” refers to the Altai, and Zhizhi Chanyu’s army by the time of reaching the Kangju was only a few thousand men strong, he was also defeated in battle and slain in this area by the Han [4].


Later on the Xiongnu split into a Northern Xiongnu and southern Xiongnu, and the Altai region was a core territory of the northern Xiongnu. It is here where the Northern Xiongnu were defeated at the battle of the Altai with the Northern Chanyu fleeing, which more or less ended their role as a significant political factor in the steppes. Soon after the remnants of the Northern Xiongnu were defeated by the Xianbei and incorporated into their empire.


Using Vahaduo Admixture JS, here are some models for these samples, keeping it simple. Since there are several streams of ancestry which can overlap I turned on the ADC function at 0.25 to reduce overfitting.


Target

Distance | ADC: 0.25x RC

MNG_North_N

CHN_Amur_River_EN

RUS_Sintashta_MLBA

CHN_Upper_Yellow_River_LN

IRN_Tepe_Hissar_C

RUS_Tyumen_HG

SBB001:SBB001.A0101

0.02480856

59.6

0.0

35.4

0.0

4.6

0.4

SBB002:SBB002.A0101

0.03926090

10.4

77.8

2.8

4.8

4.2

0.0

SBB003:SBB003.A0101

0.03166866

51.6

0.0

22.8

14.2

7.8

3.6

SBB005:SBB005.A0101

0.03431243

45.6

40.6

8.0

5.8

0.0

0.0

SBB007:SBB007.A0101

0.02634784

56.6

37.0

3.2

0.0

3.2

0.0

SBB008:SBB008.A0101

0.02331714

66.8

0.0

3.0

15.8

6.0

8.4

SBB010:SBB010.A0101

0.01981236

56.0

0.0

32.4

2.2

9.4

0.0

TAK002:TAK002.A0101

0.02331341

11.6

28.2

0.2

55.6

1.2

3.2

TAK006:TAK006.A0101

0.04395687

35.2

64.8

0.0

0.0

0.0

0.0

TAK008:TAK008.A0101

0.02917008

19.6

0.0

30.4

11.0

16.0

23.0

TAK009:TAK009.A0101

0.02572385

24.4

0.0

26.6

10.4

18.6

20.0

Average

0.02924474

39.8

22.6

15.0

10.9

6.5

5.3



Target

Distance | ADC: 0.25x RC

MNG_Slab_Grave_EIA_1

MNG_Sagly_EIA_4

CHN_Western_Liao_River_BA_o

CHN_Zaghunluq_IA

CHN_Upper_Yellow_River_IA

KAZ_Otyrar_Antiquity

SBB001:SBB001.A0101

0.02663864

5.8

70.4

23.8

0.0

0.0

0.0

SBB002:SBB002.A0101

0.04132098

88.6

0.0

9.0

0.0

0.0

2.4

SBB003:SBB003.A0101

0.03313430

32.2

51.6

0.0

3.2

13.0

0.0

SBB005:SBB005.A0101

0.02956482

77.8

9.4

12.8

0.0

0.0

0.0

SBB007:SBB007.A0101

0.02012349

62.4

0.0

31.6

0.0

0.0

6.0

SBB008:SBB008.A0101

0.02191129

67.8

0.0

0.0

25.6

6.6

0.0

SBB010:SBB010.A0101

0.03212252

0.0

69.0

22.0

3.4

5.6

0.0

TAK002:TAK002.A0101

0.01613926

53.4

0.0

0.0

2.4

44.2

0.0

TAK006:TAK006.A0101

0.02833390

34.4

0.0

65.6

0.0

0.0

0.0

TAK008:TAK008.A0101

0.03320593

0.0

62.2

0.0

22.2

0.0

15.6

TAK009:TAK009.A0101

0.02670786

0.0

43.2

0.0

47.0

0.0

9.8

Average

0.02810936

38.4

27.8

15.0

9.4

6.3

3.1



My friend Altvred who some of you might know from our collaboration “When did the Western Steppe Herder genetic profile form?” also had taken a look at the Y-DNA of the samples. Unfortunately there is no data on mtdna available at the moment so Y-DNA will have to do:


Sample

Y-DNA

Yfull clade

SBB001

C2a1a1b1

C-F1756

SBB010

C2a1a1b1

C-F1756

TAK006

C2a1a1b1

C-F1756

TAK008

R1a1a1b2a

R-Z94

TAK009

R1b2b

R-PH200



A few things to note here:


One thing I have been quite convinced of lately is that Proto-Turkic speakers originally were present in central-eastern Mongolia and the Transbaikal region during the period circa 500 BC - 100 BC. This is based on a combination of archaeology, linguistics and ancient DNA. 


I am also convinced that C-F1756 was one of the core y-chromosome haplogroup lineages amongst these early Proto-Turkic peoples (one of being key here). Now you might say “hold up, isn’t C-F1756 a Mongolic haplogroup?” and you wouldn’t be wrong as modern distribution certainly suggests that. Many cases of C-F1756 so far also have been found in samples with a Mongolic-like autosomal profile and/or in contexts such as Xianbei or Rouran sites. C-F1756 also shows up quite a bit in samples from central-eastern Mongolia during the Xiongnu period, predominantly in the area I just highlighted as what I consider the most likely location of a Proto-Turkic urheimat. And we do have samples with C-F1756 in early medieval Europe in Khazar, Avar and Hungarian conqueror period contexts which predate the presence of Mongols in Europe.


But the lineage is certainly not frequently found amongst medieval Turkic peoples. I think that there are multiple layers of y-chromosome haplogroup introgression that ultimately brought the C-F1756 frequency amongst Turks down by a significant margin, with one key event being the Xiongnu collapse and the interactions between native Altai Scythians and new eastern migrants in the ensuing periods. I am working on something regarding my opinions on the Turkic urheimat, which will cover this exact topic too but in more detail and with a lot more evidence for my claims. However it is not going to be published soon, there is still much I need to work out.


The fact that C-F1756 was quite significantly present amongst the males in this sampleset is interesting, and could indicate what I suggested earlier: C-F1756 was significantly present amongst early Turkic speaking peoples until they started mixing with local Scythian remnants around the Altai region, which lead to a replacement of C-F1756 by lineages such as R-Z93, Q1b, J2 etc. Later on as Turkic people started expanding more and more lineages introgressed into their population.


What I also wonder is if the C-F1756 samples are all related in the sense that we have sampled a dynasty or if we just have samples that arose from a common population where C-F1756 was a common lineage hence it being present at both sites. Given that Shombuuzyn Belchir doesn’t seem to be an elite site in particular, and that Han-related ancestry isn’t ubiquitous it might be the second scenario.


SB0001 and SB0010 are interesting samples because they highlight how quickly an autosomal profile can shift. For instance we have many samples with western/south siberian lineages that have a near entire east Asian profile. Here we have a sample which is over 50% Eastern Scythian related yet had C-F1756 as a paternal lineage.


Unfortunately since the article is not out yet we have no burial contexts for the samples and it is hard to figure out who is who exactly.  When it comes to Takhiltyn Khotgor, there are apparently over a hundred burials present at the site and there are several square tombs with an elite connotation. Most of the information in articles seems to be in regards to the later excavations and if the TAK samples come from samples of those excavations, I think there is decent room to speculate about the burial context of the samples at Takhyltin Khotgor.


For instance it would be interesting to note that tomb 82 and tomb 64, which are the elite square-shaped tombs, both contained women, with burial 82 being dubbed “the princess tomb” [5].


Tomb 82 of Takhiltyn Khotgor



Tomb 64 contained an older woman which had given birth before based on her pelvis remains.


Tomb 64 of Takhiltyn Khotgor


Aerial view of Tomb 64 and tomb 82 respectively.


I suspect that the TAK002 sample might come from one of these elite burials as this sample belongs to a woman, and her autosomal profile suggests a significant amount of Han-related ancestry. A legacy of the Heqin policy perhaps?


The satellite burials around tomb 82 and tomb 25 contain people buried in standard Xiongnu period graves, although there is some deviation as their burial position is flexed. The people in these tombs are described as having robust muscle attachments, an osteological sign of performing significant physical labour. The anthropological type seems to be related to populations of Manchuria, Baikal and central-eastern Mongolia. Perhaps the significantly amur shifted sample TAK006 comes from one of these burial contexts. From a genetic perspective there seems to be a connection between TAK006 and the samples at Shombuuzyn Belchir. 


Satellite burials of tomb 25 at Takhiltyn Khotgor


I wonder if TAK008 and TAK009 samples come from these aforementioned satellite burials with deviating burial rites, as the stone lining without coffins and flexed burial positions are quite reminiscent of Eastern Scythian burial rites. 


Satellite burials at tomb 64 of Takhiltyn Khotgor


That said it might be that these samples come from completely different burials. We will find out eventually when the article comes out.


From a genetic perspective TAK008 and TAK009 are two samples which are highly interesting to me because they have a  rather distinctive profile. In general you can divide “westerners” amongst the Xiongnu samples into three clusters: 


  1. Scytho-Siberian cluster.


The first one is the Scytho-Siberian cluster, which are samples with an autosomal profile similar to either Scythians from the Altai and Tian Shan regions. The steppes of the Altai mountain region were home to Scytho-Siberians during the first millennium b.c, and several of the eastern Scythian material cultures have full continuity into the first century BC, more than a century after being under Xiongnu rule. Some samples however resemble Saka from the Tian Shan region, which can be seen as a two-way mixture of Eastern Scythians and Southern Central Asians.



  1. Southern Central Asian cluster


The second cluster are samples in which the dominant factor is southern Central Asian ancestry, although there is variable complex mixture with iron age steppe nomads going on as well. The source of this ancestry primarily seems to be the Tian Shan Saka kind but some samples do seem to have Sarmatian ancestry too, which is interesting. There really is only one area where all of this comes together, and that is the Kangju state. A state founded by a nomadic population centered around Sogdia. This state was also an ally of the Xiongnu, although the relation was loose at times so we have a historic precedent for their presence amongst Xiongnu sites in Mongolia.


  1. East Xinjiang cluster


The third cluster is a rather unique one. Amongst Xiongnu period sites there is a consistent appearance of a population with partial ancestry from Scytho-Siberians, southern Central Asians, upper Yellow River inhabitants and also having quite a high amount of West-Siberian hunter-gatherer related ancestry. The samples with this profile also often had y-chromosome R1b-PH155, which was first found in the Xiaohe-Gumugou samples known for their Tarim Mummies. R1b-PH155 was also significantly present at Zaghunluq, an iron age site also known for their Tarim mummies, such as Cherchen man. The autosomal profiles of the samples in this cluster are reminiscent of those found in early iron age sites in Xinjiang such as Zaghunluq, although they can best be described as a mixture of Scytho-Siberians and Iron age Xinjiang. The most likely area for the development of this profile is the Eastern Xinjiang region as archaeology suggests the formation of a nomadic material culture of mixed Pazyryk-related and Yanbulak origin.


Shirenzigou burial site


The samples Tak008 and Tak009 clearly belong to this third cluster based on their autosomal ancestry and Y-dna ancestry. Other samples include the two “Central Asian'' Xiongnu samples  DA38 and DA41, with the latter having Y-chromosome R1b-PH155 as well. The Shirenzigou samples from the Barkol steppes fit this profile too, and are actually the reason why I dubbed it the “East Xinjiang cluster” as the earliest phases of the nomadic presence at the Shirenzigou and Heiguoliang sites predate the Xiongnu empire, and thus the presence of this cluster within Xiongnu contexts comes through the populations of the Barkol-Yiwu region.


Target

Distance | ADC: 0.25x RC

RUS_Sintashta_MLBA

RUS_Tyumen_HG

MNG_North_N

CHN_Upper_Yellow_River_LN

IRN_Tepe_Hissar_C

MNG_Xiongnu_Central_Asian:DA38

0.02588333

21.2

22.0

20.6

16.0

20.2

MNG_Xiongnu_Central_Asian:DA41

0.02477495

24.8

17.2

24.6

20.0

13.4

CHN_Shirenzigou_IA:F004

0.04137018

19.0

19.2

23.4

24.2

14.2

CHN_Shirenzigou_IA:M010

0.02621386

19.6

19.4

16.6

31.8

12.6

CHN_Shirenzigou_IA:M15-1

0.05459725

12.2

36.4

10.8

9.0

31.6

CHN_Shirenzigou_IA:M819

0.02768645

23.8

19.6

19.0

22.6

15.0

CHN_Shirenzigou_IA:M820

0.04128422

36.4

18.8

15.2

10.0

19.6

CHN_Shirenzigou_IA:M827

0.04578673

24.0

17.6

28.6

18.4

11.4

TAK008:TAK008.A0101

0.02917008

30.4

23.0

19.6

11.0

16.0

TAK009:TAK009.A0101

0.02572385

26.6

20.0

24.4

10.4

18.6

Average

0.03424909

23.8

21.3

20.3

17.3

17.3


I also suspect that these East Xinjiang cluster samples are the Yuezhi (and Wusun) based on the historical records suggesting that prior to moving to the Ili valley and Bactria the Yuezhi were present in the regions steppes of Eastern Xinjiang and Western Ganu, as well as the archaeological presence of this material culture both predating and continuing through the Xiongnu period. We know from history that several “Xiao Yuezhi” groups rather than migrating away remained under the Xiongnu, and even founded city states in modern day Xinjiang.


A few months ago I started working on a blog post which covered several of the historical peoples of the Xiongnu period that could be linked to Scythian material cultures and it is something I still haven’t finished. I currently have over 40 pages of text on this topic and honestly I am not even close to wrapping it up. I am thinking of splitting it into pieces so that it would cover several blog posts. One of them will be about the Yuezhi and their origins, and my arguments about the Yuezhi being this “East Xinjiang cluster” will be explained more thoroughly. Look at me teasing around, I didn’t mean to but these samples are hitting the topic of two WIP blog entries I have been working on at a snail’s pace.


Noin-Ula tapestry, likely depicting Kushans from Central Asia


Yeniseian-related profiles however seem to be entirely lacking amongst these samples, which troubles the interpretation that “The Xiongnu” or their elites were Yeniseian speakers. Aside from proposed loanwords and etymologies, the most secure argument for a Yeniseian presence amongst the Xiongnu is an attested phrase of the Jié people, which were one of the tribes of the Xiongnu which formed a short-lived state called Later Zhao under Shi Le during the period of Sixteen Kingdoms of the Five Barbarians. The phrase was recorded by the Kuchean monk Foutudeng, in chinese characters, glossed with a chinese translation:


秀支 替戾剛 僕谷 劬禿當

army - go out - Liu Yao's barbarian title - capture


Which according to Pulleyblank should be rendered as the following in Middle Chinese [6]:

i̯u-ci̯e - tʰei-let/lei-kɑŋ - bok/buk-kuk/yo - ɡi̯u̯o-tʰuk-tɑŋ


The Yeniseian interpretation according to Vovin [7] is as following:

“Suke t-i-r-ek-ang bok-kok k-o-t-o-kt-ang”


“Armies have gone out. [They] will catch Bokkok.”


I think the linguistic arguments behind this phrase being Yeniseian in origin are quite sensible, because it correlates with linguistic developments in Southern Yeniseian in particular [8]. This is interesting because southern Yeniseian groups such as the Arints were nomadic pastoralists living in Yurts, although their language suggests adoption of this lifestyle by way of Turkic influence, probably through the Kyrgyz [9].


The proposed Turkic etymologies from a linguistic perspective are far less sound. However, to extrapolate that the “Xiongnu language” was Yeniseian or that their elites were Yeniseians (who later language-flipped into Oghuric Turkic) based on this one group is something I find a bit baseless. It didn’t make too much sense to me to begin with because the early Xiongnu confederations were attested just north of China, and the Proto-Yeniseian homeland was considerably further north.


Empress Ashina


Last month an article about Empress Ashina was published and the genomes of her ancestry were made public yesterday.

Ancient genome of Empress Ashina reveals the Northeast Asian origin of Göktürk Khanate

Abstract:

We have unveiled the first genomic profile of the ancient Türkic royal family and Chinese historical celebrities. Our genomic analyses of Empress Ashina revealed Göktürk's Northeast Asian origin (97.7% Northeast Asian ancestry and 2.3% West Eurasian ancestry), refuting the western Eurasian origin and multiple origin hypotheses. We found Ashina shared most genetic affinity with post-Iron Age Tungusic and Mongolic Steppe pastoralists, such as Rouran, Xianbei, Khitan, and Heshui_Mohe, and showed genetic heterogeneity with other ancient Türkic people, suggesting the multiple sources of the Türkic Khanate populations. Furthermore, the limited contribution from ancient Göktürk found in modern Turkic speaking Article populations once again validates a cultural diffusion model over a demic diffusion model for the spread of Turkic languages.

Tomb of Empress Ashina. Source.


Davidski too made coordinates of her genomes:

  • A42801_scaled,0.026179,-0.41332,0.086738,-0.043928,-0.03416,-0.021196,0.00329,0.025845,-0.002454,0.012028,-0.029555,-0.01154,0.003865,-0.006881,0,0.008221,-0.002217,0.003674,-0.001257,0.02176,-0.01672,-0.00643,-0.012941,-0.005543,-0.000838
  • A42801_unscaled,0.0023,-0.0407,0.023,-0.0136,-0.0111,-0.0076,0.0014,0.0112,-0.0012,0.0066,-0.0182,-0.0077,0.0026,-0.005,0,0.0062,-0.0017,0.0029,-0.001,0.0174,-0.0134,-0.0052,-0.0105,-0.0046,-0.0007

Unfortunately the coverage of the sample isn’t all too good, so I wouldn’t take the percentages of the samples too literally, using similar models as I had used for the Xiongnu samples earlier I get something like this:


Target: A42801_scaled

Distance: 3.8806% / 0.03880631

62.4 CHN_Amur_River_EN

30.4 CHN_Upper_Yellow_River_LN

7.2 RUS_Sintashta_MLBA


Target: A42801_scaled

Distance: 4.0703% / 0.04070271

38.0 MNG_Slab_Grave_EIA_1

26.8 CHN_Western_Liao_River_BA_o

26.2 CHN_Upper_Yellow_River_IA

9.0 MNG_Sagly_EIA_4


I think it possible the authors probably underestimated the degree of western ancestry as they modeled the sample with MNG_North_N as the East Eurasian source. The issue is that the MNG_North_N samples have a small amount of ANE ancestry, which more or less lines up with the amount of Sintashta/Sagly ancestry you see pop up here.


Target: A42801_scaled

Distance: 6.5378% / 0.06537781

95.2 MNG_North_N

4.8 RUS_Sintashta_MLBA


 In any case it shows that the Early Turks of the Ashina clan at this time were undoubtedly significantly of eastern steppe origin. Furthermore they made rather weird claims about the spread of Turkic languages based on this sample:


“In summary, we have unveiled the first genomic profile of the ancient Türkic royal family. Our genomic analyses of EmpressAshina revealed Göktürk's Northeast Asian origin (97.7%Northeast Asian ancestry and 2.3% West Eurasian ancestry),refuting the western Eurasian origin and multiple origin hypotheses. We found Ashina shared most genetic affinitywith post‐Iron Age Tungusic and Mongolic Steppe pastoralists,such as Rouran, Xianbei, Khitan, and Heshui_Mohe, and showed genetic heterogeneity with other ancient Türkic people,suggesting the multiple sources of the Türkic Khanatepopulations. Furthermore, the limited contribution from ancient Göktürk found in modern Turkic‐speaking populations once again validates a cultural diffusion model over a demic diffusion model for the spread of Turkic languages.


Turkic languages did not spread due to a demic diffusion of Gokturks, Turkic languages spread due to the formation of the Gokturk empire which incorporated many Turkic peoples, by then undoubtedly having diverse genetic profiles, which then definitely had a demic diffusion across the central asian steppes. If it was merely a demic diffusion we would not see a consistent amount of significant east asian ancestry in European, Central Asian, south Siberian and Mongolian samples from Turkic contexts.


All in all, a very interesting week when it comes to ancient genetics of the steppes!

References:

  1. Miller, Bryan & Makarewicz, Cheryl & Jamsranjav, Bayarsaikhan & Tüvshinjargal, Tömörbaatar. (2018). Stone lines and burnt bones: Ritual elaborations in Xiongnu mortuary arenas of Inner Asia. Antiquity. 92. 1-19. 10.15184/aqy.2018.136.

  2. Miller (2009). Elite Xiongnu Burials at the Periphery: Tomb Complexes at Takhiltyn Khotgor, Mongolian Altai 

  3. U. Brosseder, J. Bayarsaikhan, Bryan K. Miller, Ts.Odbaatar  (2011). SEVEN RADIOCARBON DATES FOR XIONGNU BURIALS IN WESTERN AND CENTRAL MONGOLIA

  4. С.С. Сорокин (1956). О датировке и толковании Кенкольского могильника.// КСИИМК. Вып. 64. 1956.С.3-14.

  5. Bryan K. Miller, Jamsranjav Bayarsaikhan, Tseveendorj Egiimaa, Christine Lee (2007). Xiongnu Elite Tomb Complexes in the Mongolian Altai Results of the Mongol-American Hovd Archaeology Project, 2007

  6. Pulleyblank (1964). The oconsonantal system of old Chinese, part II.

  7. Vovin, Alexander (2000) "Did the Xiongnu speak a Yeniseian language?". Central Asiatic Journal 44/1 , pp. 87-104.

  8. Vovin, Alexander; Vajda, Edward J.; Vaissière, Etienne de la. (2016). "WHO WERE THE *KJET (羯) AND WHAT LANGUAGE DID THEY SPEAK?".

  9. Тимонина, Людмила Георгиевна (1984). Тюркские заимствования в енисейских языках в сравнительно-историческом освещении